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At the time of first contact, the indigenous cultures were quite different from those of the proto-industrial and mostly Christian immigrants. Some of the Northeastern and Southwestern cultures in particular were matrilineal and operated on a more collective basis than the Europeans were familiar with. The majority of Indigenous American tribes maintained their hunting grounds and agricultural lands for use of the entire tribe. Europeans at that time had patriarchal cultures and had developed concepts of individual property rights with respect to land that were extremely different. The differences in cultures between the established Native Americans and immigrant Europeans, as well as shifting alliances among different nations in times of war, caused extensive political tension, ethnic violence, and social disruption. Even before the European settlement of what is now the United States, Native Americans suffered high fatalities from contact with European diseases spread throughout the Americas by the Spanish to which they had yet not acquired immunity. Smallpox epidemics are thought to have caused the greatest loss of life for indigenous populations, although estimates of the pre-Columbian population of what today constitutes the U. S. vary significantly, from 1 million to 18 million.

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